The question is 10 female dogs average 16 puppies each year with 50% of those puppies being female. Every year the owners remove 60% of the female puppies and all of the male puppies.

What will the growth be of female puppies left and their mothers over 15years?

Only the original mothers and 20% of female puppies can have puppies (each year); the rest will be neutered.

I know this is pretty easy to work out with a calculator if you keep entering the numbers, but is there a equation that will work it out?

Thanks for any help working towards finding it.